Leadership behavior is a widespread phenomenon in animal groups on the move such as insect swarms, bird flocks, fish shoals and herds ( [1], [2], [3], [4], [5], [6], [7] ). Mathematical modeling based on the analysis of observational data is often used to study leadership behavior ( [8], [9], [10]). The rules of interaction of individuals have been inferred from observation. (For example, a zonal rule showed that the attraction and repulsion of neighbors for mosquitofish depends on the distance between them [9].) However, these proposed rules have not been tested empirically, as the live individuals are hard to control [11].